Restoring grassland on arable land: an example of a fast spontaneous succession without weed-dominated stages.

Vegetation succession was studied for 12 years on an abandoned nutrient- poor small field surrounded by a strip of natural grassland. No fertilizers or herbicides were applied. Few weeds or annuals were present during this period. Only two plants, i.e. Agrostis capillaris and Festuca rubra, dominated during the 12 years. At two spatial scales (0.04 and 0.4 m2) a dramatic increase in species richness was recorded during the first two years. However, no further trend in species richness occurred after the sixth year of succession. Successional changes were directed and continuous. The rate of succession depended on the distance from the neighbouring meadow. Vegetation at the margins of the transect developed more slowly and diverged from the middle during the succession.